General Overview
The membranous labyrinth (labyrinthus membranaceus) forms a closed system suspended inside the osseous labyrinth. Its primary feature is that it topographically mimics the contours of the bony labyrinth, yet is considerably smaller.
The entire internal cavity of this structure is filled with fluid—endolymph.
Anatomo-functionally, the membranous labyrinth is divided into several key components, each serving a distinct physiological role:
- utricle (utriculus);
- saccule (sacculus);
- membranous semicircular ducts (ductus semicirculares);
- cochlear duct (ductus cochlearis).
Vestibule: Utricle and Saccule
Two major expansions of the membranous labyrinth are located within the vestibule, responsible for vestibular functions.
The utricle (utriculus) lies in the elliptic recess (recessus ellipticus). The saccule (sacculus) is located in the spherical recess (recessus sphericus).
Specialized sensory areas called maculae are situated on the inner surface of these structures:
- macula utriculi;
- macula sacculi.
These regions contain otolith crystals known as statoconia (statoconia). Their main function is to detect linear acceleration and register changes in head position in space.
Both structures (utricle and saccule) are interconnected via the utriculosaccular duct (ductus utriculosaccularis). This duct continues as the endolymphatic duct (ductus endolymphaticus), which terminates blindly in the endolymphatic sac (saccus endolymphaticus).
Membranous Semicircular Ducts
The membranous semicircular ducts (ductus semicirculares) lie within their corresponding osseous semicircular canals.
Each of the three ducts features a characteristic dilation at one end called the membranous ampulla (ampulla membranacea). Inside the ampullae are sensory structures termed ampullary crests (crista ampullaris). Unlike the maculae of the utricle and saccule, the receptors of the ampullary crests specialize in detecting angular acceleration (rotational movements).
Cochlear Duct and Spiral Organ
The cochlear duct (ductus cochlearis) is located within the spiral canal of the cochlea (canalis spiralis cochleae). On cross-section, this duct has a distinct triangular shape.
The walls of the cochlear duct are formed by the following structures:
- Basilar membrane (membrana basilaris) — the floor/lower wall.
- Vestibular membrane or Reissner's membrane (membrana vestibularis) — the roof/upper wall.
- Outer wall — formed by the vascular stria (stria vascularis). The vascular stria is responsible for secreting (producing) the endolymph that fills the membranous labyrinth.
The basilar membrane supports the primary organ of hearing, the spiral organ (organum spirale), also known as the organ of Corti. It serves as the peripheral auditory receptor apparatus.
The spiral organ consists of:
- inner and outer hair cells (cellulae pilosae internae et externae), which act as mechanoreceptors;
- supporting cells;
- tectorial membrane (membrana tectoria), which overhangs the hair cells.